Jennifer Alice Clack, FRS (née Agnew; born 3 November 1947) is a paleontologist, who studies the "fish to tetrapod" transition— the origin, evolutionary development and radiation of early tetrapods and their relatives among the lobe-finned fishes. She is best known for her book Gaining Ground: the Origin and Early Evolution of Tetrapods, published in 2002 (second edition, 2012) and written with the layman in mind.

Clack and her team discovered the remains of the Devonian tetrapods Acanthostega and Ichthyostega in East Greenland in 1987. Acanthostega is a transitional, water-bound primitive tetrapod. Athough fragments of the skull had been discovered in 1933 by Gunnar Säve-Söderbergh and Erik Jarvik, the significance of this species as a transitional organism between fish and amphibian was deciphered by Jennifer A. Clack.

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